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macula (noun, pl. maculae) – a small pale spot or blotch on the upper or outer surface of a thallus, often due to uneven distribution of photobiont cells below the thalline cortex (or unevenly thickened cortex), representing a locally discoloured or alga-less area, not associated with any break in the cortex; usually whitish or pale; usually visible only with a lens. Related terms: reticulum, stria, striation. This is:part of upper thallus surface.

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